It is pretty well understood ___controls the flow of carbon dioxide in and out the atmosphere today填that what 还是what 为什么?

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Itisprettywellunderstood___controlstheflowofcarbondioxideinandouttheatmospheretoday填thatwhat还是what为什

It is pretty well understood ___controls the flow of carbon dioxide in and out the atmosphere today填that what 还是what 为什么?
It is pretty well understood ___controls the flow of carbon dioxide in and out the atmosphere today
填that what 还是what 为什么?

It is pretty well understood ___controls the flow of carbon dioxide in and out the atmosphere today填that what 还是what 为什么?
填that what.
因为understood后缺少宾语,that 在这里代指后面的一整句话作宾语;而后面的一整句话又缺少主语,用what.