英语翻译To what extent competition determines the composition of a community and the density of particular species has been the source of considerable controversy.感觉怎么有两个动词?determines 和has been,what从句到哪截止?

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英语翻译Towhatextentcompetitiondeterminesthecompositionofacommunityandthedensityofparticularspecieshasbe

英语翻译To what extent competition determines the composition of a community and the density of particular species has been the source of considerable controversy.感觉怎么有两个动词?determines 和has been,what从句到哪截止?
英语翻译
To what extent competition determines the composition of a community and the density of particular species has been the source of considerable controversy.
感觉怎么有两个动词?determines 和has been,what从句到哪截止?

英语翻译To what extent competition determines the composition of a community and the density of particular species has been the source of considerable controversy.感觉怎么有两个动词?determines 和has been,what从句到哪截止?
(生存)竞争对于群落构成和特定物种密集度究竟有多大的决定性曾引发大规模的论战
has been 谓语 后面是宾语部分
to what extent 是一起的,别把他们分开,按中文习惯句子就像这样
competition determines (the composition of a community and the density of particular species) To what extent (has been.)

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